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Why is the pocket between beta subunits in the R state of heme smaller?

2 confomers of HB

T and R

T is more stable (lower affinity to O2) b/c its has greater ion pairs and another thing I noticed is that is has larger ‘pocket’ between beta subunits

 

Why would the R state have a smaller pocket bewteen b subunits? (That does not seem conducive to having a higher affinity to O2)

 

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